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hampes-threadwort-cephaloziella-hampeana.jpg from: https://www.earth.com/plant-encyclopedia/Bryophytes/Cephaloziellaceae/cephaloziella-hampeana/en/

Cephaloziella triplicata: A Tiny But Mighty Moss

Introduction

When it comes to the world of mosses, Cephaloziella triplicata S.W.Arnell may be tiny in size, but it is a fascinating species that plays important ecological roles. This diminutive moss, a member of the

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Cephaloziella_dentata_009C.JPG from: https://cisfbr.org.uk/Bryo/Cornish_Bryophytes_Cephaloziella_dentata.html

Cephaloziellaceae family, is commonly known simply as Cephaloziella

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2020-09-17-14-12-24.jpg from: https://www.britishbryologicalsociety.org.uk/learning/species-finder/cephaloziella-divaricata/

. Let’s take a closer look at this marvelous miniature plant.

Background

Cephaloziella triplicata is a species of leafy liverwort, which are non-vascular plants in the division Marchantiophyta, class Jungermanniopsida. The species epithet “triplicata” refers to the trilobed leaves that are a key identification feature. It was first described scientifically in 1956 by Swedish botanist Sigfrid Wilhelm Arnell.

Morphology and Identification

Cephaloziella triplicata forms dense mats of tiny green shoots only 1-2 mm tall

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Cephaloziella_stellulifera_012.JPG from: https://cisfbr.org.uk/Bryo/Cornish_Bryophytes_Cephaloziella_stellulifera.html

. The minuscule leaves are only 0.2-0.4 mm long and are divided into 3 lobes

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medium.jpeg from: https://www.inaturalist.org/taxa/56495-Cephaloziella-divaricata

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cephaloziella-divaricata-a-144443-481819-4278-M0TJT2.jpg from: https://www.alamy.com/stock-photo-cephaloziella-divaricata-a-144443-481819-4278-172645314.html

(hence “triplicata”). Leaves are succubously inserted at a 45° angle.

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cephaloziella_divaricata.jpg from: https://wnmu.edu/academic/nspages/gilaflora/cephaloziella_divaricata.html

Oil bodies, organelles unique to liverworts, are found in all leaf cells. The underleaves are small or absent.
Cephaloziella can be a tricky genus to identify to species, often requiring microscopic examination. However, the distinctly trilobed leaves and obtuse lobe tips help distinguish C. triplicata from similar species like C. varians and C. hampeana. The small size and dense growth also separate it from other genera like Cephalozia.

Global Distribution and Habitat

Cephaloziella triplicata has a

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3327-l-3.jpg from: https://www.wildflowers.co.il/hebrew/picture.asp?ID=19936

widespread but scattered distribution, being found in:

It grows on soil, rocks, rotting logs, and tree bases in forests and other shaded, damp habitats from lowlands to subalpine elevations. It is often found in disturbed sites.

Ecological Roles and Adaptations

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CephaloziellaDivaricata5862Habit.jpg from: https://lichens.twinferntech.net/pnw/species/Moss_or_liverwort_species.shtml

As a pioneer species, C. triplicata plays an important role in soil formation and stabilization. The dense mats help retain moisture and build up organic matter. This allows other plants to colonize and succession to occur.
The small size is an adaptation that allows C. triplicata to exploit cracks and crevices that other plants cannot grow in. The leaves’ lobed structure increases surface area for water and nutrient uptake. Oil bodies may help with desiccation tolerance.
C. triplicata also provides food and shelter for invertebrates. Mites, springtails, and other tiny organisms inhabit the miniature “forests” created by moss mats.

Conclusion

Cephaloziella triplicata may be an easily overlooked inhabitant of the forest floor, but it serves essential pioneering and engineering roles in ecosystems worldwide. Next time you’re in the woods, take a moment to appreciate the mighty miniature world of mosses beneath your feet. What other secrets of the small scale remain to be uncovered?