
better+cross+timmiella+crassinervis.jpg from: https://mosswalks.blogspot.com/2013/07/moss-mites-and-timmiela-crassinervus.html
Timmiella subintegra Dixon: The Fascinating Moss of the Timmiellaceae Family
Introduction
Mosses are often overlooked, but they play crucial roles in ecosystems around the world. One particularly interesting moss is Timmiella subintegra Dixon, also known simply as Timmiella. This small but mighty plant is part of the Timmiellaceae family in the Bryophyta phylum and Bryopsida

medium.jpg from: https://enciclovida.mx/especies/147289-timmiella-anomala
class. Let’s take a closer look at this fascinating moss species.
Background
Timmiella subintegra Dixon was first described by the British bryologist Hugh Neville Dixon in 1930. It is found in various regions across the globe, typically growing on soil, rocks, or tree bases in dry habitats. As a moss, Timmiella lacks true roots, stems, and leaves like vascular plants. Instead, it has simple structures that allow it to photosynthesize, absorb water and nutrients, and reproduce.

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Morphology and Identification
Timmiella subintegra forms small, dense cushions or mats. Its shoots are usually under 1 cm tall. The leaves are lanceolate (lance-shaped) and have a

large.jpeg from: https://www.naturalista.mx/guide_taxa/1398185
prominent costa (midrib) that often extends beyond the leaf tip. Leaf margins are entire to slightly toothed near the apex. Timmiella is dioicous, meaning male and female reproductive structures are on separate plants.
Global Distribution and Habitat
This moss has a wide distribution, being found in Europe, Africa, Asia, Australia, and the Americas

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. It grows in dry, exposed habitats like rock outcrops, soil banks, and at the bases of trees. Timmiella tolerates disturbance and low nutrient availability, often being a pioneer species in early successional environments.
Ecological Roles and Adaptations

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As with other mosses, Timmiella plays important roles in its ecosystems:

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- Helps retain moisture and prevent soil erosion
- Provides habitat for micro-organisms and small invertebrates
- Pioneers in disturbed areas and enriches soil as it breaks down
- Indicator of air quality as it absorbs pollutants
Timmiella has adaptations that allow it to thrive in dry conditions:
- Papillose leaf cells that help trap moisture
- Ability to dry out and rehydrate quickly
- Rhizoids that anchor it to substrates and absorb water and nutrients

0116.jpeg from: https://www.calflora.org/app/taxon?crn=12935

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Characteristic | Description |
---|---|
Size | Small cushions or mats, shoots < 1 cm tall |
Leaves | Lanceolate with prominent costa, entire to toothed margins |
Habitat | Dry exposed sites on soil, rock, tree bases |
Distribution | Europe, Africa, Asia, Australia, Americas |
Reproduction
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Dioicous |
Adaptations | Papillose cells, desiccation tolerance, rhizoids |
Conclusion
Timmiella subintegra Dixon may be small, but it is a remarkable moss with a wide distribution and important ecological roles. Its ability to thrive in challenging conditions makes it a valuable pioneer species. Next time you’re out in nature, take a closer look – you might just spot some Timmiella making its mark on the world! What other mighty mosses have you encountered?