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2020-09-28-11-42-31.jpg from: https://www.britishbryologicalsociety.org.uk/learning/species-finder/philonotis-capillaris/

Exploring the Fascinating World of Philonotis capillata (Mitt.) Paris Moss

Mosses are often overlooked, but they play crucial roles in ecosystems around the world. One particularly interesting species is Philonotis capillata (Mitt.) Paris, a moss in the Bartramiaceae family. Also known simply as

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275720.jpg from: https://inpn.mnhn.fr/espece/cd_nom/4992

Philonotis, this tiny plant is worth taking a closer look at. In this blog post, we’ll dive into the details of Philonotis capillata and discover what makes it so special.

Background on Philonotis Mosses

The genus Philonotis contains around 150 species of mosses found across the globe. They belong to the division

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275721.jpg from: https://inpn.mnhn.fr/espece/cd_nom/4992/tab/carte

Bryophyta and class Bryopsida. Philonotis mosses are generally small, growing in dense tufts or cushions. Many species prefer damp or wet habitats like stream banks, seeps, and damp soil or rock.

Morphology and Identification of Philonotis capillata

Philonotis capillata is a small moss, typically growing 1-3 cm tall. The stems are slender and branched, often forming dense tufts. The leaves are lanceolate (lance-shaped) and have a distinctly toothed margin near the apex. When dry, the leaves become twisted and contorted.
One key identification feature is the presence of

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3258-l-4.jpg from: http://www.wildflowers.co.il/hebrew/picture.asp?ID=19288

papillose leaf cells

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3258-l-2.jpg from: http://www.wildflowers.co.il/hebrew/picture.asp?ID=19286

. Under a microscope, the cell surface appears bumpy or roughened. The scientific name “capillata” means “hairy” and refers to the fine, hair-like appearance of the dry leaves.

Global Distribution and Habitat

Philonotis capillata has a wide distribution, being found in many parts of the world including Europe, Asia, Africa, and the Americas. It grows in a variety of habitats, but is most commonly found on damp soil or rock along streams, rivers, and seepage slopes. The moss is able to tolerate periodic submersion and benefits from the consistent moisture.

Ecological Roles and Adaptations

Like other mosses, Philonotis capillata plays important roles in its ecosystem:

  • Erosion control: The dense tufts help stabilize soil and prevent erosion along stream banks.
  • Water retention: Moss clumps act like sponges, absorbing and slowly releasing water, regulating moisture in the microenvironment.
  • Habitat for microorganisms: The nooks and crannies among moss stems provide habitat for diverse communities of tiny organisms like rotifers, nematodes, and tardigrades.

Philonotis capillata has several adaptations that allow it to thrive in its preferred habitat:

  • Papillose leaf cells help trap and hold a film of water on the leaf surface, enabling the moss to absorb moisture and nutrients.
  • Twisted leaves contort and cling together when dry, reducing water loss. When moistened, they quickly untwist to resume photosynthesis.
  • Rhizoidal tomentum (a woolly covering of reddish filaments) helps anchor the moss to its substrate and absorb water and minerals.

In Summary

Philonotis capillata is a fascinating moss with a wide distribution and important ecological roles. Its small size belies its outsized impact in stabilizing soils, regulating moisture, and providing microhabitats in riparian ecosystems. Next time you’re near a stream, take a moment to appreciate the tiny but mighty Philonotis growing there!
What other overlooked or underappreciated mosses have you encountered? Learning more about these humble plants can give us a window into the captivating world thriving all around us, just beneath our notice.